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With each weapon level up you will recieve a skill point for that weapon that can be used to learn new skills, or improve old ones. Some skills will require a specific level to unlock.
Press K while playing to see your skills and notice the skill points listed in the lower right corner. Then mouse over the skill circles to see which skills you can "spend" those points on to improve or learn. Some will most likely be locked, but you will see what level they unlock at this way.
Be sure to poke around on the forums to learn which upgrades you should use and when. Careful use of skill points will help you advance while careless use could actually make some things more difficult.
Good luck mate!
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